Know more

About cookies

What is a "cookie"?

A "cookie" is a piece of information, usually small and identified by a name, which may be sent to your browser by a website you are visiting. Your web browser will store it for a period of time, and send it back to the web server each time you log on again.

Different types of cookies are placed on the sites:

  • Cookies strictly necessary for the proper functioning of the site
  • Cookies deposited by third party sites to improve the interactivity of the site, to collect statistics

Learn more about cookies and how they work

The different types of cookies used on this site

Cookies strictly necessary for the site to function

These cookies allow the main services of the site to function optimally. You can technically block them using your browser settings but your experience on the site may be degraded.

Furthermore, you have the possibility of opposing the use of audience measurement tracers strictly necessary for the functioning and current administration of the website in the cookie management window accessible via the link located in the footer of the site.

Technical cookies

Name of the cookie


Shelf life

CAS and PHP session cookies

Login credentials, session security



Saving your cookie consent choices

12 months

Audience measurement cookies (AT Internet)

Name of the cookie


Shelf life


Trace the visitor's route in order to establish visit statistics.

13 months


Store the anonymous ID of the visitor who starts the first time he visits the site

13 months


Identify the numbers (unique identifiers of a site) seen by the visitor and store the visitor's identifiers.

13 months

About the AT Internet audience measurement tool :

AT Internet's audience measurement tool Analytics is deployed on this site in order to obtain information on visitors' navigation and to improve its use.

The French data protection authority (CNIL) has granted an exemption to AT Internet's Web Analytics cookie. This tool is thus exempt from the collection of the Internet user's consent with regard to the deposit of analytics cookies. However, you can refuse the deposit of these cookies via the cookie management panel.

Good to know:

  • The data collected are not cross-checked with other processing operations
  • The deposited cookie is only used to produce anonymous statistics
  • The cookie does not allow the user's navigation on other sites to be tracked.

Third party cookies to improve the interactivity of the site

This site relies on certain services provided by third parties which allow :

  • to offer interactive content;
  • improve usability and facilitate the sharing of content on social networks;
  • view videos and animated presentations directly on our website;
  • protect form entries from robots;
  • monitor the performance of the site.

These third parties will collect and use your browsing data for their own purposes.

How to accept or reject cookies

When you start browsing an eZpublish site, the appearance of the "cookies" banner allows you to accept or refuse all the cookies we use. This banner will be displayed as long as you have not made a choice, even if you are browsing on another page of the site.

You can change your choices at any time by clicking on the "Cookie Management" link.

You can manage these cookies in your browser. Here are the procedures to follow: Firefox; Chrome; Explorer; Safari; Opera

For more information about the cookies we use, you can contact INRAE's Data Protection Officer by email at or by post at :


24, chemin de Borde Rouge -Auzeville - CS52627 31326 Castanet Tolosan cedex - France

Last update: May 2021

Menu Logo Principal AgroParisTech Université de Lorraine UMR SILVA

Home page

COLIN Francis

Senior researcher (« Directeur de recherches’) Co-leader of the research team PHARE (25 permanent persons + 5 PhD students) Leader of the projects LABEX « BARK-TAN-BIO » and MAAF-FEDER « EXTRA FOR EST
Francis Colin

COLIN Francis

Research Unit: SILVA

Research Team: PHARE ‘Physiology of the tree in its environment’
Centre INRAE Grand Est -Nancy
F-54280 CHAMPENOUX (France)

+33 (0) 3 83 39 40 42

+33 (0) 7 82 17 05 98


2003: Accreditation for supervising researches (HDR): Nancy University
1992: PhD in Forest Science: Engref-Nancy (now AgroParisTech-Nancy)
1990: Forest Engineer: Engref-Nancy (now AgroParisTech-Nancy) 
1988: Master in Forest Science: Nancy University

Scientific activities

  • Themes

♦ Chemical wood properties especially extractives contents

♦ Tree development and architecture - Branching (forks, sequentials and epicormics)

♦ Biomass estimation of trunk, branches, bark, heartwood, juvenile wood, knots

♦ Wood formation in branches and knots

♦Silviculture and wood quality

  • Measurement instruments

♦ X-ray tomography

♦ Terrestrial Lidar

♦ Network of experimental plots

♦ Statistical modelling

  • PhD theses: Supervisor (S) and Co-supervisor (CoS):

♦ Rodolphe Bauer (2017-2020) (S). Estimation of the volume of bark, heartwood and knots. Within-tree and interspecific variability (MAAF et FEDER Lorraine)

♦ Antoine Billard (2017-2020) (S). Analysis of the within-tree and interspecific variability of wood density (ADEME – Région Grand-Est)

♦ Van-Tho Nguyen (2015-2018) (S). Estimation of the quality with T-Lidar. In collaboration with T. Constant (LERFoB), A. Piboule (ONF RDI Nancy), A. Krähenbühl, B. Kerautret, I. Debled-Rennesson (LORIA)

♦ Jialin Song (2014-2017) (S) QTL analysis of the branching of Quercus robur. In collaboration with Antoine Kremer, Christophe Plomion, Catherine Bodénès (UMR BIOGECO Bordeaux) et Oliver Brendel (UMR EEF Nancy)

♦ Zineb Kebbi-Benkeder (2012-2015) CoS)Interspecific and intraspecific biodiversity of knot extractives. AgroParisTech PhD in Wood science and fibres. 186p.  In collaboration with P. Gérardin et S. Dumarçay (LERMAB)

♦ Jean-Baptiste Morisset, (2009-2012) (S). X-Raytomography ; analysis and modelling of the ontogenesis of the epicormics of Quercus petraea (L.) Matt.). Thèse AgroParisTech PhD in Forest Sciences. 142p  and annexes. In collaboration with F. Mothe (LERFoB)

  • Research management


       ► French projects

2017-2020. Projet MAAF – FEDER Lorraine « EXRAFOR-EST » (ressources régionales en biomasse ligno-cellulosique pour une utilisation en chimie des extractibles)

2017-2019. LABEX « Arbre » Projet innovant « BARK-TAN-BIO » (variabilité intra-arbre et interspécifique des taux d’extractibles dans l’écorce)

2016. LABEX « Arbre » Projet « PROTEXTRA » (mesure du duramen des branches) (Responsable)

2016. LABEX « Arbre » Projet « Post-doc Zineb » (Post-doc à OSU Corvallis de Z. Kebbi-Benkeder ) (Responsable)

2015. LABEX « Arbre » Projet AFE « OAK-Epic » (Effet site sur la détermination des QTLs de ramification du chêne pédonculé) (Responsable)

2014. LABEX « Arbre » Projet AFE « Menzies0 »  (accueil de Nate Osborne de OSU Corvallis) (Responsable)

2012-2015. Contrat ONF-INRA ModelFor3 (Responsable d’actions) Actions « Génétique des épicormiques du chêne » et « Qualité du Sapin et du Douglas »

2012-2015. LABEX “Arbre”  Projet « Wood acclimation in disturbed environment » (« WADE »);  Responsable de tâche « Branch Wood »

2011. Contrat ONF-INRA ModelFor2. Responsable de thème 3 : « Croissance et qualité des bois » 

2005-2010. Contrat ONF-INRA ModelFor2. Responsable d’action A31 : « Élaboration de la qualité commerciale des grumes de Hêtre et de Chêne »

2004-2005. Contrat ONF-INRA 4839B. Analyse des situations sylvicoles à risque de présence élevée de gourmands dans les peuplements de chêne sessile et pédonculé (Responsable)

       ► European projects

1997-1998 : FAIR-BM96.2554 Recommendations for afforestation of agricultural set-aside with oak species (Coordinateur)

1996-1999 : OAK-KEY (PL95-0823) New silvicultural alternatives in young oak high forests. Consequences on high quality timber production (Coordinateur)

1994-1998. Root-Arch  (CT93-1269) Aspects of sustainability by afforestation of agricultural set aside areas : development of roots and shoot/root ratios (Resp. INRA)

1992-1994. FOREST (CT91-0024) Silvicultural control and non-destructive assessment of timber quality in plantation grown spruces and Douglas fir (Resp. Tâche)

  • Conference organization

International conference « EPIC-CONF » Nancy (France) September 14-17 2017 (chairman)

International IUFRO « MeMoWood » conference ; group 5.01.04 « Wood Quality Modelling »  Nancy (France) October 1-4 2013 (chairman)

  • Extension

♦ Silviculture and improvment of wood products products

♦ Forest resources estimation (solid wood, extractives…)

♦ Continuing education: silviculture and wood quality


(AgroParisTech Nancy & Master AETPF ‘Agrosciences, Environnement, Territoires, Paysage, Forêt’ Université de Lorraine)

♦ Biology of tree development

♦ Silviculture and wood properties

♦ Storm damages in forests

Scientific papers

  • Kebbi-Benkeder Z., Manso R., Dumarçay S., Gérardin P., Chopard B., Colin F., 2017. Knot extractives: a model for analyzing the eco-physiological factors that control the within and between-tree variability. Tree Physiology 31 (5) : 1619–1633 doi:10.1007/s00468-017-1573-z
  • §  Song J., Brendel O., Bodénès C., Plomion C., Kremer A., Colin F., 2017. X-ray computed tomography to decipher the genetic architecture of tree branching traits: oak as a case study. Tree Genetics and Genomes 13 (5): 1-15 doi: 10.1007/s11295-016-1083-y

All publications

All Francis Colin publications
Read more